Ecological Archives M075-010-A2

T. Royama. 2005. Moran effect on nonlinear population processes. Ecological Monographs 75:277–293.

Appendix B. Autocorrelation for u’t, an estimator of ut by the moving-average method. A pdf file of this appendix is also available.

Figure A2 (see Appendix A) shows the sample autocorrelation functions, ruu(l) and ru’u’(l), for the density-independent influence ut and its estimator u’t (Fig. 6b). The ruu(l), as samples of ruu(l) º 0, l  > 0, are insignificantly different from zero at all lags shown, whereas the ru’u’(l) exhibit a significant negative spike at lag 2, which can be explained as follows.

Consider that the series {Rt} is made up of two components: a low-frequency cycle (gt, say) due to the autocorrelated density-dependent process and a rapid fluctuation (about the cycle) due to the density-independent influence ut with no autocorrelation. Thus, we may write

Rt = gt + ut.      
(A.2)

After the moving-average transformation on both sides of (A2), we have

R’t = g’t + u”t     
(A.3)

in which the single and double primes indicate the moving-average transformations of the respective letters in Eq. A.2. Recalling the definition u’t = Rt - R’t in the main text, the difference (A.2) - (A.3) yields

gt  - g’t + ut - u”t  = u’t.

Now that, by assumption, gt is void of the rapidly fluctuating component, the difference

|gt - g’t| would be small compared with |ut - u”t (= zt, say)| such that u’t would be approximately equal to zt. Hence, letting k (an odd positive integer) be the order of the moving-average transformation, the autocorrelation function ru’u’(l) would be approximated by rzz(l) whose theoretical function is readily shown to be

rzz(l) = - (k + l)/k(k - 1), 
0 < l £ (k - 1)/2
 
     
= (k - l)/k(k - 1),
(k - 1)/2 < l < k
 
     
= 0, 
k < l.
(A.4)

 For k = 5, rzz(1) = - 0.3, rzz(2) = - 0.35, rzz(3) = 0.1, rzz(4) = 0.05, and rzz(l ³ 5) = 0, as in the ru’u’(l) based on the length 50, although for length 4000 only the ru’u’(2) is distinct. The tendency for ru’u’(2) to be distinctly negative shows up in the cross-correlation function ru’v’(l) (Fig. 7, middle column) that tends to exhibit significant spikes at lags other than 0.

The results (A.4) show that rzz(l) tends to zero for all l as the order k increases, apparently suggesting that the higher the order k, the better ru’v’(l) approximates ruv(l). This is true when gt is linear, or flat, across all t. Otherwise, g’t would over-smooth gt sooner or later as k increases, resulting in the difference (gt  - g’t) to increase in variance. Conversely, too low an order results in an under-smoothing that also results in an increase in variance. Thus, there must be an optimal order at which the variance of the difference (gt  - g’t) is minimized. Delving further into this subject is beyond the present scope. However, a graphical inspection (cf. Fig. 6a) would probably be adequate in practice.



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